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Understanding the intricately interconnected systems that regulate chromosome biology and the cell cycle is a major challenge in modern cell biology. Yeast systems continue to provide key biological insights and technical breakthroughs that drive progress in the fields of cell cycle, DNA replication, recombination and repair, chromatin structure, spindle assembly and regulation, kinetochore structure and function, telomere biology and nuclear organization. This FASEB SRC brings together leaders in these fields, young up-and-coming faculty and the students and postdocs, who are actually doing the work to share the most current technical and conceptual advances, meet their colleagues from across the globe and establish new collaborations.