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The International Symposium on Hepatitis C Virus and Related Viruses has been a recurring annual conference since 1994. It has become the main forum for basic science investigators working on hepatitis C virus (HCV) and related viruses. Over the past 25 years, this conference has been instrumental in developing and advancing major discoveries in the HCV field including the understanding of the virus life cycle and development of in vitro replication systems and animal models that led to the development of direct acting antivirals (DAA) that have revolutionized HCV treatment.