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Despite the dramatic recent advances in cell biology and evolutionary biology, remarkably little interaction exists between the two fields. Because all evolutionary change ultimately begins with alterations at the cellular level, this deficiency represents a major gap in biological understanding. The goal of this program is to assemble a diverse set of scientists - from cell biology, evolutionary biology, biophysics, biochemistry, and applied mathematics - with interdisciplinary interests, in a setting that will facilitate long-lasting collaborative research and the establishment of a formal field of evolutionary cell biology.