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A number of recent discoveries have radically changed the picture of gene and chromatin regulation, as system-level organisational mechanisms have emerged to play a key role. This meeting, now in its third edition, aims to highlight new biological breakthroughs in such an important research area and the crucial contribution from quantitative approaches, both in experiment and modelling. It will provide a unique opportunity to bring together researchers working in such a vast, yet strategic, field in disciplines ranging from biology and medicine to chemistry, computer science, engineering, mathematics and physics.