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An unprecedented number of new pathogenic fungi and variants of extant strains are emerging to cause disease in animals and plants, putting the resilience of wild and managed ecosystems in jeopardy. This meeting will unite researchers sharing a common aim – to exploit advances in biology to understand the drivers causing the emergence of fungi and to forge a research agenda to mitigate their impact. This meeting is free to attend, but advance registration via the meeting webpage is essential. This meeting will be followed by a related satellite meeting 'Tackling emerging fungal threats to animal health, food security and ecosystem resilience: the state of the art' on 9 - 10 March 2016 at the Royal Society at Chicheley Hall, home of the Kavli Royal Society Centre, Buckinghamshire.