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This FASEB Science Research Conference will integrate viral genomics, molecular virology, biophysical virology, and structural virology. Invited speakers will showcase diverse approaches to study virus assembly and structure, viral replication, entry into host cells (infection) the interactions and co-evolution between viruses and hosts, and the deep connections between viruses from all kingdoms of life. The meeting will address recent developments in the area of structure determination, such as in single particle electron-microscopic image reconstruction, tomography and asymmetric reconstruction where advances continue to be rapid and exciting. The diversity of the scientific program and the ample opportunity for interactions among attendees encourages the development of new cross-disciplinary collaborations.