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Mitochondria and chloroplasts differ from other cellular compartments by their endosymbiotic origin, their semiautonomous genetic machineries and their much more complex biology. This GRC brings together established scientists and young researchers to report on and discuss emerging aspects of organelle biology. It has a strong focus on basic biological processes and an interdisciplinary nature that brings together scientists from the plant, animal and fungal communities. Specialized sessions are dedicated to the evolution of organelle-containing eukaryotes, the gene expression systems of organelles, the import of proteins and assembly of organellar complexes from proteins, lipids and cofactors, organelle-associated signaling pathways, large-scale and systems biology approaches, the interactions between and dynamics of organelles, and the removal of organelles by autophagy during aging and senescence. For all these topics, leading experts will report on novel exciting discoveries and developments. For each session of the GRC on Mitochondria and Chloroplasts, an introduction by a Discussion Leader will provide a general overview to integrate the research communities and stimulate conceptual discussions. Speakers with international recognition in the field will join young investigators and scientists who have not previously attended this conference. Short talks will be selected from abstracts to showcase emerging late-breaking topics, especially from early-career investigators. Daily poster sessions will further enrich and stimulate interactions between participants and research communities. This GRC is preceded by a GRS.